for his work on ribonucleaseandfor their contribution to the understanding of the connection between chemical structure and catalytic activity of the ribonuclease molecule
for their elucidation of the enzymatic mechanism underlying the synthesis of adenosine triphosphateandfor his discovery of an ion-transporting enzyme, Na+, K+ -ATPase
... are held about twice as long as their short counterparts. Also, there exists a third, extra-long length for most vowels, which is used in various cases, but particularly ...